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Prof. Dr. Bettina Reichenbacher


Gierl, C., Liebl, D., Šanda, R., Vukić, J., Esmaeili, H.R., Reichenbacher, B. (in press). What can goby otolith morphology tell us? Cybium.

Janssen, R., Doppler, G., Grimm, K., Grimm, M., Haas, U., Hiss, M., Köthe, A., Radtke, G., Reichenbacher, B., Salamon, M., Standke, G., Teipel, U., Thomas, M., Uffenorde, H., Wieland-Schuster, U. & Subkommission Tertiär-Stratigraphie (2018). Das Tertiär in der Stratigraphischen Tabelle von Deutschland 2016. [The Tertiary in the Stratigraphic Table of Germany 2016 (STG 2016).] – Z. Dt. Ges. Geowiss, 169 (2), p. 267–294. Link to article.

Jolivet, M., Barrier, L., Dauteuil, O., Laborde, A., Li, Q., Reichenbacher, B., Popescu, S., Sha, J., Guo, Z. (2018). Late Cretaceous - Palaeogene topography of the Chinese Tian Shan: New insights from geomorphology and sedimentology. Earth and Planetary Science Letters, 499: 95–106. Link to article.

Kevrekidis, C., Valtl, M., Penk, S.B.R., Altner, M. & Reichenbacher, B. (2018). Rebekkachromis nov. gen. from the middle-upper Miocene (11 MYA) of Central Kenya: The oldest record of a haplotilapiine cichlid fish. Hydrobiologia, online first, 1–26. doi: 10.1007/s10750-018-3715-8

Pippèrr, M. & Reichenbacher, B., Kirscher, U., Sant, K, Hanebeck, H. (2018). The middle Burdigalian in the North Alpine Foreland Basin (Bavaria, SE Germany) – a lithostratigraphic, biostratigraphic and magnetostratigraphic re-evaluation. Newsletter Stratigraphy, 51 (3): 285–309. Link to article.

Reichenbacher, B., Filipescu, S., Miclea, A. (2018). A unique middle Miocene fish fauna from coastal deposits in the southeastern Pannonian Basin (Romania). Palaeodiversity and Palaeoenvironment, online first, doi: 10.1007/s12549-018-0334-3

Reichenbacher, B., Gregorová, R., Holcová, K., Šanda, R., Vukić, J., Přikryl, T. (2018). Discovery of the oldest Gobius (Teleostei, Gobiiformes) from a marine ecosystem of Early Miocene age. Journal of Systematic Palaeontology, 16(6): 493-513. Link to article.

Teimori, A., Esmaeili, H. R., Hamidan, N., & Reichenbacher, B. (2018). Systematics and historical biogeography of the Aphanius dispar species group (Teleostei: Aphaniidae) and description of a new species from Southern Iran. Journal of Zoological Systematics and Evolutionary Research. Online first, doi:10.1111/jzs.12228.

Vahed, N.S., Esmaeili, H.R., Masoudi, M. & Reichenbacher, B. (2018). Early otolith development in the critically endangered tooth-carp Aphanius farsicus (Teleostei: Cyprinodontidae). Environmental Biology of Fishes, 101:1309–1317. doi: 10.1007/s10641-018-0778-3. Link to article.


Altner, M., Schliewen, U., Penk, S.B.R., Reichenbacher, B. (2017). † Tugenchromis pickfordi gen. et sp. nov. from the upper Miocene – a stem-group cichlid of the ‘East African Radiation’. Journal of Vertebrate Paleontology, 37(2): e1297819. Click here to download this article (open access).

Gierl, C. & Reichenbacher, B. (2017). Revision of so-called Pomatoschistus (Gobiiformes, Teleostei) from the Late 
Eocene and Early Oligocene. Palaeontologia Electronica 20.2.33A: 1-17. Click here to download this article (open access).

Hofmann, F., Reichenbacher, B., Farley, K.A. (2017). Evidence for >5 Ma paleo-exposure of an Eocene-Miocene paleosol of the Bohnerz Formation, Switzerland. Earth and Planetary Science Letters, 465: 168–175.

Pippèrr, M. & Reichenbacher, B. (2017). Late Early Miocene palaeoenvironmental changes in the North Alpine Foreland Basin. Palaeogeography, Palaeoclimatology, Palaeoecology, 468, 485–502.

Rasmussen, C., Reichenbacher, B., Lenz, O., Altner, M., Penk, S.B.R., Prieto, J., Brüsch, D. (2017). Middle–late Miocene palaeoenvironments, palynological data and a fossil fish Lagerstätte from the Central Kenya Rift (East Africa). Geological Magazine, 154(1): 24–56.

Sant, K., Kirscher, U., Reichenbacher, B., Pippèrr, M., Jung, D., Doppler, G. & Krijgsman, W. (2017). Late Burdigalian sea retreat from the North Alpine Foreland Basin: New magnetostratigraphic age constraints. Global and Planetary Change, 152: 38–50.

Teschner, C. & Reichenbacher, B. (2017). Otolith-based age determination of mid-Burdigalian marine sediments in the North Alpine Foreland Basin. Bulletin of Geosciences 92(2): 143–152.  Click here to download this article (open access).


Ghanbarifardi, M., Esmaeili, H.R., Gholami, Z., Aliabadian, M., Reichenbacher, B. (2016). Molecular phylogeny of three mudskippers (Gobiidae) from the Persian Gulf and Gulf of Oman (Makran). Journal Applied Ichthyology, 32, 416–420.

Masoudi, M., Esmaeili, H.R., Teimori, A., Gholami, Z., Gholamhosseini, A., Sayyadzadeh, G., Keivany, Y., Reichenbacher, B. (2016). Sympatry and possible hybridization among species of the killifish genus Aphanius Nardo, 1827 (Teleostei: Cyprinodontidae) in Southwestern Iran. Limnologica, 59 (2016): 10–20.

Pippèrr, M., Reichenbacher, B., Doppler, G., Hagmaier, M. & Jung, D. (2016). The northern coast of the Ottnangian (middle Burdigalian, early Miocene) Molasse Sea in Germany: sediments, foraminiferal assemblages and biostratigraphy. International Journal of Earth Sciences, 105: 1055–1085.

Reichenbacher, B. (2016). Jean Gaudant (* 31 December 1939 — † 6 December 2015) In Memoriam. Geodiversitas 38, 327-339. Click here to download this article (open access).

Salvador, R.B., Pippèrr, M., Reichenbacher, B., Rasser, M.W. (2016). Early Miocene continental gastropods from new localities of the Molasse Basin in southern Germany. Paläontologische Zeitschrift, 90(3): 469-491.


Altner, M. & Reichenbacher, B. (2015). †Kenyaichthyidae fam. nov. and †Kenyaichthys gen. nov. – First Record of a Fossil Aplocheiloid Killifish (Teleostei, Cyprinodontiformes). PLoS ONE 10(4):e0123056. Click here  to download this article (open access).

Bratishko, A., Schwarzhans, W., Reichenbacher, B., Vernyhorova, Y., Ćorić, S. (2015): Fish otoliths from the Konkian (Miocene, early Serravallian) of Mangyshlak (Kazakhstan): testimony to an early endemic evolution in the Eastern Paratethys. Paläontologische Zeitschrift, 89: 839–889.

Gaudant, J., Barrón, E., Anadón, P., Reichenbacher, B. & Peñalver, E. (2015). Palaeoenvironmental analysis of the Miocene Arcas del Villar gypsum sequence (Spain), based on palynomorphs and cyprinodontiform fishes. Neues Jahrbuch Geologie, Paläontologie, Abhandlungen, 277(1): 105–124.

Gholami, Z., Esmaeili, H.R., Reichenbacher, B. (2015). New data on the zoogeography of Aphanius sophiae (Teleostei: Cyprinodontidae) in the Central Zagros (Southwest Iran). Limnologica, 51: 70–82.

Gholami, Z., Esmaeili, H.R., Erpenbeck, D., Reichenbacher, B. (2015). Genetic connectivity and phenotypic plasticity in the cyprinodont Aphanius farsicus from the Maharlu Basin, South-western Iran. Journal of Fish Biology, 86: 882–906.

Gierl, C. , Reichenbacher, B. (2015): A new Fossil Genus of Gobiiformes From the Miocene Characterized by a Mosaic Set of Characters. Copeia, 103(4): 792–805.

Jost, J., Kälin, D., Börner, S., Vasilyan, D., Lawver, D., Reichenbacher, B. (2015). Vertebrate microfossils from the Upper Freshwater Molasse in the Swiss Molasse Basin: Implications for the evolution of the North Alpine Foreland Basin during the Miocene Climate Optimum. Palaeogeography, Palaeoclimatology, Palaeoecology, 426: 22–33.


Esmaeili, H.R., Teimori, A., Sayyadzadeh, G., Masoudi, M., Reichenbacher, B. (2014). Phylogenetic relationships of the tooth-carp Aphanius (Teleostei: Cyprinodontidae) in the river systems of southern and south-western Iran based on mtDNA sequences. Zoology in the Middle East, 60:1, 29-38.

Esmaeili, H.R., Teimori, A., Gholami, Z., Reichenbacher, B. (2014). Two new species of the tooth-carp Aphanius (Teleostei: Cyprinodontidae) and the evolutionary history of the Iranian inland and inland-related Aphanius species. Zootaxa 3786(3): 246-268.

Gholami, Z., Esmaeili, H.R., Erpenbeck, D., Reichenbacher, B. (2014). Phylogenetic analysis of Aphanius from the endorheic Kor River Basin in the Zagros Mountains, Southwestern Iran (Teleostei: Cyprinodontiformes: Cyprinodontidae). Journal of Zoological Systematics and Evolutionary Research, 52(2), 130-141.

Reichenbacher, B. & Reichard, M. (2014). Otoliths of Five Extant Species of the Annual Killifish Nothobranchius from the East African Savannah. PLoS ONE 9(11): e112459. Click here to download this article (open access).

Teimori, A., Esmaeili, H.R., Erpenbeck, D., Reichenbacher, B. (2014). A new and unique species of the genus Aphanius (Teleostei: Cyprinodontidae) from Southern Iran: A case of regressive evolution. Zoologischer Anzeiger, 253, 327–337.


Annabi, A., Said, K., Reichenbacher, B. (2013). Inter-population differences in otolith morphology are genetically encoded in Aphanius fasciatus (Cyprinodontiformes, killifishes). Scientia Marina, 77(2): 269–279. pdf (2,6 MB)

Becker, D., Marty, D., Pirkenseer, C., Cavin, L., Grimm, K., Reichenbacher, B. (2013). A tribute to the late Professor Jean-Pierre Berger (8 July 1956–18 January 2012). Swiss Journal of Geosciences 106: 119–124.

Gholami, Z., Teimori, A., Esmaeili, H.R., Schulz-Mirbach, T., Reichenbacher, B. (2013). Scale surface microstructure and scale size in the tooth-carp genus Aphanius (Teleostei, Cyprinodontidae) from endorheic basins in Southwest Iran. Zootaxa 3619 (4): 467–490.

Gierl, C., Reichenbacher, B., Gaudant, J., Erpenbeck, D. & Pharisat, A. (2013). An Extraordinary Gobioid Fish Fossil from Southern France. PLoS ONE 8(5): e64117. Click here to download this article.

Pirkenseer, C., Berger, J.-P., Reichenbacher, B. (2013). The position of the Rupelian/Chattian boundary in the southern Upper Rhine Graben based on new records of microfossils. Swiss Journal of Geosciences 106: 291–301.

Reichenbacher, B., Krijgsman, W., Lataster, Y., Pippèrr, M., van Baak, C.G.C., Chang, L., Kälin, D., Jost, J., Doppler, G., Jung, D., Prieto, J., Abdul Aziz, H., Böhme, M., Garnish, J., Kirscher, U., Bachtadse, V. (2013). An alternative magnetostratigraphic framework for the Lower Miocene (Burdigalian / Ottnangian, Karpatian) in the North Alpine Foreland Basin. Swiss Journal of Geosciences 106: 309–334.


Esmaeili, H.R., Teimori, A., Gholami, Z., Zarei, N., Reichenbacher, B. (2012). Re-validation and re-description of an endemic and threatened species, Aphanius pluristriatus (Jenkins, 1910) (Teleostei, Cyprinodontidae), from southern Iran. Zootaxa, 3208: 58–67.

Teimori, A., Schulz-Mirbach, T., Esmaeili, H.R. & Reichenbacher, B. (2012a). Geographic differentiation of Aphanius dispar (Teleostei: Cyprinodontidae) from Southern Iran. Journal of Zoological Systematics and Evolutionary Research 50(4): 289-304.

Teimori, A., Esmaeili, H.R., Gholami, Z., Zarei, N., Reichenbacher, B. (2012b). Aphanius arakensis, a new species of tooth-carp (Cyprinodontidae) from the endorheic Namak Lake basin in Iran. Zookeys, 215: 55-76.

Teimori, A., Jawad, L.A.J., Al-Kkarusi, L.H., Al-Mamry, J.N., Reichenbacher, B. (2012c). Late Pleistocene to Holocene diversification and zoogeography of the Arabian killifish Aphanius dispar inferred from otolith morphology. Scientia Marina, 76(4): 637-645. [Featured Article] pdf (2,2 MB)


Carnevale, G., Haghfarshi, E., Abbasi, S., Alimohammadian, H. & Reichenbacher, B. (2011). A new species of silverside (Teleostei: Atheriniformes) from the Late Miocene of NW-Iran. Acta Palaeontologica Polonica, 56: 749-756.

Reichenbacher, B., Alimohammadian, H., Sabouri, J., Haghfarshi, E., Faridi, M., Abbasi, S., Matzke-Karasz, R., Fellin, M.G., Carnevale, G., Schiller, W., Vasilyan D. & Scharrer, S. (2011). Late Miocene stratigraphy, palaeoecology and palaeogeography of the Tabriz Basin (NW Iran, Eastern Paratethys). Palaeogeography, Palaeoclimatology, Palaeoecology, 311: 1-8.

Teimori, A., Esmaeili, H.R. & Reichenbacher, B. (2011). Aphanius farsicus, a replacement name for A. persicus (Jenkins, 1910) (Teleostei, Cyprinodontidae). Zootaxa, 3096: 53–58. pdf (1,2 MB)

Schneider, S., Pippèrr, M., Frieling, D. & Reichenbacher, B. (2011). Sedimentary facies and paleontology of the Ottnangian Upper Marine Molasse and Upper Brackish Water Molasse of eastern Bavaria: A field trip guide. In: Carena, S., Friedrich, A.M. & Lammerer, B. (Eds.): Geological Field Trips in Central Western Europe: Fragile Earth International Conference, Munich, September 2011: Geological Society of America, Field Guide 22, p. 35–50.


Heckeberg, N., Pippèrr, M., Läuchli, B., Heimann, F. & Reichenbacher, B. (2010). The Upper Marine Molasse (Burdigalian, Ottnangian) in southwestern Germany – facies interpretation and a new lithostratigraphic terminology. Zeitschrift deutsche Gesellschaft für Geowissenschaften, 161: 285-302.

Pippèrr, M. & Reichenbacher, B. (2010). Foraminifera from the borehole Altdorf (SE Germany): proxies for Ottnangian (Early Miocene) palaeoenvironments of the Central Paratethys. Palaeogeography, Palaeoclimatology, Palaeoecology, 289: 62-80.


Frieling, D., Pippèrr, M., Schneider, S. & Reichenbacher, B. (2009a). Sedimentology and stratigraphy at the rocky coast of the upper Burdigalian Molasse Sea: a case study from Gurlarn near Passau (SE Germany). Facies, 55:47–62.

Frieling, D., Mazumder, R. & Reichenbacher, B. (2009b). Tidal sediments in the Upper Marine Molasse (OMM) of the Allgäu area (Lower Miocene, Southwest-Germany). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen, 254: 151-163.

Heimann, F. U. M., Schmid, D. U., Pippèrr, M. & Reichenbacher, B. (2009). Re-interpreting the Baltringer Horizont as a subtidal channel facies: Implications for a new understanding of the Upper Marine Molasse "Cycles" (Early Miocene). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen, 254: 135-149.

Pippèrr, M. & Reichenbacher, B. (2009. Biostratigraphy and paleoecology of benthic foraminifera from the Eggenburgian "Ortenburger Meeressande" of southeastern Germany (Early Miocene, Paratethys). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen, 254: 41-61.

Reichenbacher, B. & Kowalke, T. (2009). Neogene and present-day zoogeography of killifishes (Aphanius and Aphanolebias) in the Mediterranean and Paratethys areas . Palaeogeography, Palaeoclimatology, Palaeoecology, 281: 43-56.

Reichenbacher, B., Feulner, G.R. & Schulz-Mirbach, T. (2009a). Geographic Variation in Otolith Morphology Among Freshwater Populations of Aphanius dispar (Teleostei, Cyprinodontiformes) from the Southeastern Arabian Peninsula. – Journal of Morphology, 270: 469–484.

Reichenbacher, B., Kamrani, E., Esmaeili, H.R. & Teimori, A. (2009b). The endangered cyprinodont Aphanius ginaonis (Holly, 1929) from southern Iran is a valid species: evidence from otolith morphology. Environmental Biology of Fishes, 86: 507-521.

Vasilyan, D., Reichenbacher, B., Carnevale G. (2009). A new fossil Aphanius species from the Upper Miocene of Armenia (Eastern Paratethys). Paläontolgische Zeitschrift, 83: 511-519.


Schulz-Mirbach, T. & Reichenbacher, B. (2008). Fossil Aphanius (Teleostei, Cyprinodontiformes) from southwestern Anatolia (Turkey): A contribution to the evolutionary history of a hotspot of freshwater biodiversity. Geodiversitas, 30(3): 577-592.

Schulz-Mirbach, T., Stransky, C., Schlickeisen, J. & Reichenbacher, B. (2008). Differences in otolith morphologies between surface- and cave-dwelling populations of Poecilia mexicana (Teleostei, Poeciliidae) reflect adaptations to life in an extreme habitat. Evolutionary Ecology Research, 10: 537–558.

Schulz-Mirbach, T., Scherb, H. & Reichenbacher, B. (2008). Are hybridization and polyploidization phenomena detectable in the fossil record? – A case study on otoliths of a natural hybrid, Poecilia formosa (Teleostei: Poeciliidae). Neues Jahrbuch Geologie Paläontologie Abhandlungen, 249(2): 223-238.

Weidmann, M., Berger, Engesser, B., Reichenbacher, B., Sauvagnat, J. & Schäfer, P. (2008). La Molasse de la Vallée de Joux (Jura, Suisse et France). [The Joux Valley molasse (Jura, Switzerland and France)]. Bulletin de la Société Vaudoise des Sciences Naturelles, 91/1: 69-101.


Charollais, J. Weidmann, M., Berger, J.P., Engesser, B., Hotellier, J.F., Gorin, G., Reichenbacher, B., Schäfer, P. (2007). La Molasse du bassin franco-genevois et son substratum [The Molasse in the Greater Geneva area and its substratum]. Archives des Sciences , 60 (2-3): 59-173

Martini, E. & Reichenbacher, B. (2007). Nanoplankton und Fisch-Otolithen in der Mittleren Pechelbronn-Formation (Unter-Oligozän, Oberrheingraben/Mainzer Becken). Geologische Abhandlungen Hessen, 116: 235-273.

Pippèrr, M., Reichenbacher, B., Witt, W., Rocholl, A. (2007). The Middle and Upper Ottnangian of the Simssee area (SE Germany): Micropalaeontology, biostratigraphy and chronostratigraphy. Neues Jahrbuch Geologie Paläontologie Abhandlungen, 245: 353-378.

Reichenbacher, B., Gaudant, J. & Griessemer, T.W. (2007a). A late Burdigalien Gobiid fish, Gobius brevis (Agassiz, 1839), in the Upper Hydrobia Beds at Edenkoben in the middle Upper Rhine Graben (W-Germany). Paläontologische Zeitschrift 81(4).

Reichenbacher, B., Sienknecht, U., Küchenhoff, H. & Fenske, N. (2007b). Combined Otolith Morphology and Morphometry for Assessing Taxonomy and Diversity in Fossil and Extant Killifish (Aphanius, †Prolebias). Journal of Morphology, 268: 898–915.

Reichenbacher, B., Böhme, M. & Lang, L. (2007c). Fossilien aus der Oberen Süßwassermolasse bei Ried (Kreis Neuburg an der Donau). Archaeopteryx, 25: 49-53.

Reischenbacher, D., Rifelj, H., Sachsenhofer, R.F., Jelen, B., Ćorić, S., Gross, M. & Reichenbacher, B. (2007). Early Badenian paleoenvironment in the Lavanttal Basin (Mühldorf Beds; Austria): Evidence from paleontology and geochemistry. Austrian Journal of Earth Sciences, 100, 202-229.


Jost, J., Kälin, D., Schulz-Mirbach, T. & Reichenbacher, B. (2006). Late Early Miocene lake deposits near Mauensee, central Switzerland: fish fauna (otoliths, teeth), accompanying biota and palaeoecology. Eclogae Geologica Helvetiae, 99: 309-326.

Reichenbacher, B. & Prieto, J. (2006). Lacustrine fish faunas (Teleostei) from the Karpatian of the northern Alpine Molasse Basin, with a description of two new species of Prolebias Sauvage. Palaeontographica A, 278(1-6): 87-95.

Schulz-Mirbach, T. & Reichenbacher, B. (2006). Reconstruction of Oligocene and Neogene freshwater fish faunas - an actualistic study on cypriniform otoliths. Acta Palaeontologica Polonica, 51(2): 283-304.

Schulz-Mirbach, T. & Reichenbacher, B., Yildirim, Z. & Atalay, A. (2006). Otolith characteristics of species, subspecies and populations of Aphanius Nardo, 1827 (Teleostei, Cyprinodontiformes) from Anatolia (Turkey). Journal of Natural History, 40/27-28: 1687-1705.


Berger, J.-P., Reichenbacher, B., Becker, D., Grimm, M., Grimm, K., Picot, L., Storni, A., Pirkenseer, C., Derer, C. & Schaefer, A. (2005a). Paleogeography of the Upper Rhine Graben (URG) and the Swiss Molasse Basin (SMB) from Eocene to Pliocene. International Journal of Earth Sciences, 94(4): 697- 710.

Berger, J.-P., Reichenbacher, B., Becker, D., Grimm, M., Grimm, K., Picot, L., Storni, A., Pirkenseer, C. & Schaefer, A. (2005b). Eocene-Pliocene time scale and stratigraphy of the Upper Rhine Graben (URG) and the Swiss Molasse Basin (SMB). International Journal of Earth Sciences, 94(4): 711-731.

Doppler, G., Heissig, K. & Reichenbacher, B. (2005). Zur Gliederung des Tertiärs im süddeutschen Molassebecken. Newsletters on Stratigraphy, 41(1-3): 359-375.

Gaudant, J. & Reichenbacher, B. (2005). Hemitrichas stapfi n. sp.: a new Atherinid species (Teleostei) with otoliths in situ from the quarry "Am Katzenrech" in the Mainz Basin (Upper Oligocene). Zitteliana A, 45: 189-198.

Kowalke, T. & Reichenbacher, B. (2005). Early Miocene (Ottnangian) Mollusca of the Western Paratethys - ontogenetic strategies and palaeoenvironments. Geobios, 38(5): 609-635.

Reichenbacher, B. (2005). Die Fischfauna (Otolithen) der Unteren Süßwassermolasse von La Chaux, S. 106-108. - In: Schäfer, P.: Beiträge zur Ostracoden- und Foraminiferen-Fauna der Unteren Süßwassermolasse in der Schweiz und in Savoyen (Frankreich). 2*. La Chaux (Kanton Waadt, Schweiz). Senckenbergiana lethaea 85 (1) 95-117.

Reichenbacher, B., Kälin D. & Jost, J. (2005). A fourth St. Gallen Formation cycle (?) in the Karpatian Upper Marine Molasse of central Switzerland. Fazies, 51: 160-172.


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