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  1. Altner M & Reichenbacher B (2021) A small cichlid species flock from the Upper Miocene (9–10 MYA) of Central Kenya. Hydrobiologia 848:3613–3637. DOI: 10.1007/s10750-020-04358-z
  2. Breedy, Odalisca, Hector M. Guzman, Catalina Murillo-Cruz, and Sergio Vargas. 2021. “Discovery of a New Species of Leptogorgia Milne Edwards & Haime, 1857 (Anthozoa: Octocorallia: Gorgoniidae) from Eastern Tropical Pacific Mesophotic Reefs.” Marine Biodiversity: A Journal of the Senckenberg Research Institute / Senckenberg Forschungsinstitut Und Naturmuseum 51 (6): 95.
  3. Charmpila EA, Wagner M & Reichenbacher B (2021) First documentation of the otoliths of the species of Gouania (Teleostei: Gobiesocidae) in the Mediterranean Sea. Journal Fish Biology 98:1450–1455. DOI: 10.1111/jfb.14646
  4. Conci, Nicola, Sergio Vargas, and Gert Wörheide. 2021. “The Biology and Evolution of Calcite and Aragonite Mineralization in Octocorallia.” Frontiers in Ecology and Evolution 9: 81.
  5. Coskun OK, Vuillemin A, Schubotz F, Klein F, Sichel SE, Eisenreich W, Orsi W* (2021) Quantifying the effects of hydrogen on carbon assimilation in a seafloor microbial community associated with ultramafic rocks. The ISME Journal 16: 257-271.
  6. E Capo, C Giguet-Covex, A Rouillard, K Nota, P D Heintzman, A Vuillemin, D Ariztegui, F Arnaud, S Belle, S Bertilsson, C Bigler, R Bindler, AG Brown, CL Clarke, SE Crump, D Debroas, G Englund, GF Ficetola, RE Garner, J Gauthier, I Gregory-Eaves, L Heinecke, U Herzschuh, A Ibrahim, V Kisand, KH Kjær, Y Lammers, J Littlefair, E Messager, ME Monchamp, F Olajos, W Orsi, MW Pedersen, DP Rijal, J Rydberg, T Spanbauer, KR Stoof-Leichsenring, P Taberlet, L Talas, C Thomas, DA Walsh, Y Wang, E Willerslev, A van Woerkom, H H Zimmermann, MJL Coolen, LS Epp, I Domaizon, IG Alsos, L Parducci (2021) Lake sedimentary DNA research on past terrestrial and aquatic biodiversity: Overview and recommendations. Quarternary 4: 6.
  7. Ebert M, Bapst L, Kölbl-Ebert M, Malvesy T, López-Arbarello A (2021) Coccolepis Agassiz, 1843 (Coccolepididae, Chondrostei) from the Upper Jurassic Solnhofen Archipelago–rediscovery of the type specimen and open questions. Neues Jahrbuch für Geologie und Paläontologie-Abhandlungen 300/1: 11-22. DOI: 10.1127/njgpa/2021/0976
  8. Echreshavi S, Esmaeili HR, Teimori A & Reichenbacher B (2021) A comparative study on the caudal skeleton of goatfishes (Teleostei: Perciformes: Mullidae) from the Western Indo-Pacific region: An additional taxonomic tool. Regional Studies in Marine Science 48 (2021) 102066. DOI: 10.1016/j.rsma.2021.102066
  9. Filek T, Hofmayer F, Feichtinger I, Berning B, Pollerspöck J, Zwicker J, Smrzka D, Peckmann J, Kranner M, Mandic O, Reichenbacher B, Kroh A, Uchman A, Roetzel R & Harzhauser M (2021) Environmental conditions during the late Oligocene transgression in the North Alpine Foreland Basin (Eferding Formation, Egerian) – a multidisciplinary approach. Palaeogeography, Palaeoclimatology, Palaeoecology 580:110527. DOI: 10.1016/j.palaeo.2021.110527
  10. Forte G, Kustatscher E, Van Konijnenburg-van Cittert JHA (2021) Conifer diversity in the Middle Triassic: new data from the Fossillagerstätte Kühwiesenkopf/Monte Prà della Vacca (Pelsonian, Anisian) in the Dolomites (Northeastern Italy). International Journal of Plant Sciences 128(6): 445–467. DOI: 10.1086/714280
  11. Fürsich FT, Schneider S, Werner W, Lopez-Mir B & Pierce CP (2021) Life at the continental–marine interface: palaeoenvironments and biota of the Alcobaça Formation (Late Jurassic, Central Portugal), with a formal definition of the unit appended. Palaeobiodiversity and Palaeoenvironments. DOI: 10.1007/s12549-021-00496-x
  12. Galitz A, Nakao Y, Schupp PJ, Wörheide G, Erpenbeck D (2021) A soft spot for chemistry–current taxonomic and evolutionary implications of sponge secondary metabolite distribution. Marine Drugs 19: 448. DOI:
  13. Garber AI, Ramirez GA, McAllister SM, Orsi W, D’Hondt S (2021) Cryptic metabolisms in anoxic subseafloor sediment. Environmental Microbiology Reports 13: 696-701. DOI: 10.1111/1758-2229.12983
  14. García Massini JL, Guido DM, Campbell KC, Sagasti AJ & Krings M (2021) Filamentous cyanobacteria and associated microorganisms, structurally preserved in a Late Jurassic chert from Patagonia, Argentina. Journal of South American Earth Sciences 107:103111. DOI:
  15. Guzmán-Sandoval JA & Rössner GE (2021) Miocene chevrotains (Mammalia, Artiodactyla, Tragulidae) from Pakistan. Historical Biology 33(6): 743-776. DOI: 10.1080/08912963.2019.1661405
  16. Harper CJ & Krings M (2021) Fungi as parasites: a conspectus of the fossil record. In: de Beats K & Huntley JW (eds.), The Evolution and Fossil Record of Parasitism – Identification and Macroevolution of Parasites (Topics in Geobiology, vol. 49), pp. 69–108, Cham, Springer. DOI:
  17. Hausmann IM, Nützel A, Roden VJ & Reich M (2021) Palaeoecology of tropical marine assemblages from the Late Triassic of Misurina, Dolomites, Italy. Acta Palaeontologica Polonica 66:143-192. DOI:10.4202/app.00659.2019
  18. Höhna S, Landis MJ and Huelsenbeck JP (2021) Parallel power posterior analyses for fast computation of marginal likelihoods in phylogenetics. PeerJ, 9:e12438. DOI: 10.7717/peerj.12438
  19. Holwerda FM, Rauhut OWM, Pol D (2021) Osteological revision of the holotype of the Middle Jurassic sauropod dinosaur Patagosaurus fariasi (Sauropoda: Cetiosauridae) Bonaparte, 1979. Geodiversitas 43 (16): 575-643. DOI:
  20. Hooper JNA, Wörheide G, Hajdu E, Erpenbeck D, Voogd NJ, Klautau M (2021) Zootaxa 20 years: Phylum Porifera. Zootaxa 4979: 038-056. DOI:
  21. Karapunar B, Nützel A (2021) Slit-band gastropods (Pleurotomariida) from the Upper Triassic St. Cassian Formation and their diversity dynamics in the Triassic. Zootaxa 5042 (1): 1-165. DOI:
  22. Karapunar B, Werner W, Fürsich FT & Nützel A (2021) The earliest example of sexual dimorphism in bivalves – evidence from the astartid Nicaniella (Lower Jurassic, southern Germany. Journal of Paleontology, 95: 1216–1225. DOI: 10.1017/jpa.2021.48
  23. Karapunar B, Werner W, Fürsich FT, Nützel A (2021) The earliest example of sexual dimorphism in bivalves - evidence from the astartid Nicaniella (Lower Jurassic, southern Germany). Journal of Paleontology 95(6): 1216-1225. DOI:
  24. Ketwetsuriya C, Hausmann IM, Nützel A (2021) Diversity patterns of Middle Permian gastropod assemblages from the Tak Fa Formation, Central Thailand. Palaeobiodiversity and Palaeoenvironments 101: 907-925. DOI: 10.1007/s12549-021-00482-3
  25. Kevrekidis C, Arratia G, Bacharidis N & Reichenbacher B (2021) A new clupeid fish from the upper Miocene of Greece: A possible Hilsa relative from the Mediterranean. Acta Palaeontologica Polonica 66:3, 605–621. DOI: 10.4202/app.00871.2020
  26. Kölbl-Ebert M (2021) The Physical Sciences. In: Bullivant S & Ruse M (2021) The Cambridge History of Atheism. 2 vols., Cambridge UK: Cambridge University Press: 366–387.
  27. Krings M (2021) Peculiar bundles and a knot of thin filaments in microbial mats from the Lower Devonian Rhynie and Windyfield cherts. Review of Palaeobotany and Palynology 291:104442. DOI:
  28. Krings M (2021) Rhyniotaxillus minutulus n. sp., a pico-sized colonial cyanobacterium from the 410-million-yr-old Windyfield chert of Scotland. Nova Hedwigia 113:17–31. DOI: 10.1127/nova_hedwigia/2021/0638
  29. Krings M (2021) Stigonema (Nostocales, Cyanobacteria) in the Rhynie chert (Lower Devonian, Scotland). Review of Palaeobotany and Palynology 295:104505. DOI:
  30. Krings M (2021) The Rhynie chert land plant Aglaophyton majus harbored cyanobacteria in necrotic local lesions. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 300:279–289. DOI: 10.1127/njgpa/2021/0990
  31. Krings M (2021) Triskelia scotlandica, an enigmatic Rhynie chert microfossil revisited PalZ 95:1–15. DOI: 10.1007/s12542-020-00531-w
  32. Krings M, Serbet SM & Harper CJ (2021) Rhizophydites matryoshkae gen. et sp. nov. (fossil Chytridiomycota) on spores of the early land plant Horneophyton lignieri from the Lower Devonian Rhynie chert. International Journal of Plant Sciences 182:109–122. DOI:
  33. López-Arbarello A, Ebert M (2021) Diversity of chondrostean fish Coccolepis from the Late Jurassic Solnhofen Archipelago, Southern Germany. Acta Palaeontologica Polonica 66(4): 837-846. DOI: 10.4202/app.00873.2021
  34. Manconi R, Erpenbeck D, Fromont J, Wörheide G, Pronzato R (2022) Discovery of the freshwater sponge genus Corvospongilla Annandale (Porifera: Spongillida) in Australia with the description of a new species and phylogeographic implications. Limnology 23: 73-87. DOI:
  35. Mazaherijohari M, Kustatscher E, Roghi G, Ghasemi-Nejad E, Gianolla P (2021) A monotypic stand of Neocalamites iranensis n. sp. from the Carnian Pluvial Episode (Late Triassic) of the Aghdarband area, NE Iran (Turan Plate). Rivista Italiana di Paleontologia e Stratigrafia 127(2): 189–209. DOI: 10.13130/2039-4942/15646
  36. Mennecart B, Métais G, Costeur L, Ginsburg L & Rössner GE (2021) Reassessment of the Miocene genus Amphimoschus Bourgeois, 1873 (Mammalia, Artiodactyla, Ruminantia, Pecora). PloS ONE 16(1): e0244661. DOI: 10.1371/journal.pone.0244661
  37. Mennecart B, Métais G, Tissier J, Rössner GE & Costeur L (2021) 3D models related to the publication: Reassessment of the Miocene genus Amphimoschus Bourgeois, 1873 (Mammalia, Artiodactyla, Ruminantia, Pecora). Morpho Museum Journal 6:e131. DOI: 10.18563/journal.m3.131
  38. Moisan P, Krings M, Voigt S & Kerp H (2021) Fossil roots with root nodules from the Madygen Formation (Ladinian–Carnian; Triassic) of Kyrgyzstan. Geobios 64:65–75. DOI:
  39. Musser, Jacob M., Klaske J. Schippers, Michael Nickel, Giulia Mizzon, Andrea B. Kohn, Constantin Pape, Paolo Ronchi, Sergio Vargas, et al. 2021. “Profiling Cellular Diversity in Sponges Informs Animal Cell Type and Nervous System Evolution.” Science 374 (6568): 717–23. DOI: 10.1126/science.abj2949
  40. Nützel A (2021) Gastropods as parasites and carnivorous grazers - a major guild in marine ecosystems. In: De Baets K, Huntley J (Eds) The Evolution and Fossil Record of Parasitism: Identification and Macroevolution of Parasitism, Topics in Geobiology 49, 209-229. DOI: 10.1007/978-3-030-42484-8_6
  41. Opluštil S, Pšenička J, Šimůnek Z, Simonetto L, Kustatscher E, Votočková Frojdová, J (2021) The Late Pennsylvanian flora of the Italian Carnic Alps. Review of Palaeobotany and Palynology 285: 104360. DOI: 10.1016/j.revpalbo.2020.104360
  42. Orsi W*, Vuillemin, Coskun OK, Rodriguez P, Oertel Y, Niggemann J, Mohrholz V, Gomes-Saez G (2021) Carbon assimilating fungi from surface ocean to subseafloor revealed by coupled phylogenetic and stable isotope analysis. The ISME Journal.
  43. Orsi WD, Magritsch T, Vargas S, Coskun ÖK, Vuillemin A, Höhna S, Wörheide G, D’Hondt S, Shapiro BJ, Carini P (2021) Genome evolution in bacteria isolated from million-year-old subseafloor sediment. mBio, 12:e01150–21. DOI: DOI: 10.1128/mBio.01150-21
  44. Ortiz González, I.C., Rivera-Vicéns, R.E., Schizas, N.V., 2021. Description of four Millepora spp. transcriptomes and their potential to delimit the Caribbean fire coral species. Marine Genomics 59, 100863.
  45. Prieto J, Rummel M & Rosina VV (2021) First record of the microtoid rodent Microtocricetus Fahlbusch & Mayr 1975 from the South German karsts. Historical Biology, 1-5. DOI:10.1080/08912963.2021.2017914
  46. Puzenat V, Escartin J, Martel JE, Barreyre T, Le Moine S, Paraskevi N, Gracias N, Allemand P, Antoniou V, Coskun O, Garcia R, Grandjean P, Jørgensen SL, Magi L, Mandalakis M, Orsi W, Polymenakou P, Schouw A, Vallicrosa G, Vlasopoulos O. (2021) Shallow-water hydrothermalism at Milos (Greece): Nature, distribution, heat fluxes and impact on ecosystems. Marine Geology 438; 106521. DOI: 10.1016/j.margeo.2021.106521
  47. Rauhut OWM, Carballido JL, Pol D (2021) First osteological record of a stegosaur (Dinosauria, Ornithischia) from the Late Jurassic of South America. Journal of Vertebrate Paleontology 40 (6): e1862133. DOI:
  48. Rauhut OWM, Pol D (2021) New theropod remains from the Late Jurassic Cañadón Calcáreo Formation of Chubut, Argentina. South American Journal of Earth Sciences 111: 103434. DOI 10.1016/j.jsames.2021.103434
  49. Reich M (2021) The first Cretaceous ophiopluteus skeleton. Journal of Paleontology 95: 1284-1292. DOI: 10.1017/jpa.2021.63
  50. Reiswig, H.M., Dohrmann, M., Kelly, M., Mills, S., Schupp, P.J., Wörheide, G., 2021. Rossellid glass sponges (Porifera: Hexactinellida) from New Zealand waters, with description of one new genus and six new species. ZooKeys 1060, 33–84. DOI: 10.3897/zookeys.1060.63307
  51. Rössner GE, Costeur L & Scheyer MT (2021) Antiquity and fundamental processes of the antler cycle in Cervidae (Mammalia). The Science of Nature 108:3. DOI: 10.1007/s00114-020-01713-x
  52. Sabatini F, Pandeli E, Kustatscher E (2021) Carboniferous plant fossils from the San Lorenzo Schists (Pisani Mountains, Tuscany, Italy): a preliminary study of the palaeobotanical collection of the Museo Naturalistico Archeologico dell’Appennino Pistoiese. Rivista Italiana di Paleontologia e Stratigrafia 127(2): 397–425. DOI: 10.13130/2039-4942/15843
  53. Sadeghi R, Esmaeili HR, Teimori A, Ebrahimi M & Gholamhosseini A, Reichenbacher B, (2021) Comparative ultrastructure and ornamentation characteristics of scales in gobiid species (Teleostei: Gobiidae) using the scanning electron microscope. Microscopy Research and Technique 84:1243-1256. DOI:10.1002/jemt.23683
  54. Sadeghi R, Esmaeili HR, Zarei F, Reichenbacher B (2021) Matrilineal evidence for genetic structure and Late Pleistocene demographic expansion of the Ornate goby Istigobius ornatus (Teleostei: Gobiidae) in the Persian Gulf and Oman Sea. Marine Ecology 42:e12629. DOI: 10.1111/maec.12629#mce_temp_url#
  55. Schilling A-M & Rössner GE (2021) New skull material of Pleistocene dwarf deer from Crete (Greece). Comptes Rendus Palevol 20(9): 141-164. DOI: 10.5852/cr-palevol2021v20a9
  56. Schmidt-Schultz TH, Reich M & Schultz M (2021) Exceptionally preserved extracellular bone matrix proteins from the late Neogene proboscidean Anancus (Mammalia: Proboscidea). PalZ 95: 757-765. DOI: 10.1007/s12542-021-00566-7
  57. Schoenemann B, Clarkson ENK, Bartels C, Südkamp W, Rössner GE & Ryck U (2021) A 390 million-year-old hyper-compound eye - the visual system of phacopid trilobites. Scientific Reports 11: 19505. DOI: 10.1038/s41598-021-98740-z
  58. Schuster A, Pomponi SA, Pisera A, Cárdenas P, Kelly M, Wörheide G, Erpenbeck D (2021) Systematics of ‘lithistid’ tetractinellid demosponges from the Tropical Western Atlantic-implications for phylodiversity and bathymetric distribution. PeerJ 9: e10775. DOI:
  59. Van Konijnenburg-van Cittert JHA, Pott C, Schmeißner S, Dütsch G, Kustatscher E (2021) The Rhaetian flora of Wüstenwelsberg, Bavaria, Germany: Description of selected gymnosperms (Ginkgoales, Cycadales, Coniferales) together with an ecological assessment of the locally prevailing vegetation. Review of Palaeobotany and Palynology 288: 104398. DOI: 10.1016/j.revpalbo.2021.104398
  60. van Konijnenburg-van Cittert JHA, Schmeißner S, Dütsch G, Kustatscher E, Pott, C (2021) Plant macrofossils from the Rhaetian of Einberg near Coburg (Bavaria, Germany). Part 1. Ferns, fern allies and seed ferns. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 302(3): 331–353. DOI: 10.1127/njgpa/2021/1034
  61. Van Soest, R.W.M., Aryasari, R., Voogd, N.J., 2021. Mycale species of the tropical Indo-West Pacific (Porifera, Demospongiae, Poecilosclerida). Zootaxa 4912 (1), 1–212.
  62. Vargas, Sergio, Laura Leiva, and Gert Wörheide. 2021. “Short-Term Exposure to High-Temperature Water Causes a Shift in the Microbiome of the Common Aquarium Sponge Lendenfeldia Chondrodes.” Microbial Ecology 81 (1): 213–22.
  63. Vargas, Sergio, Thorsten Zimmer, Nicola Conci, Martin Lehmann, and Gert Wörheide. 2021. “Transcriptional Response of the Calcification and Stress Response Toolkits in an Octocoral under Heat and pH Stress.” Molecular Ecology, November. DOI:
  64. Villa A, Montie R, Röper M, Rothgänger M, Rauhut OWM (2021) Sphenofontis velserae gen. et sp. nov., a new rhynchocephalian from the Late Jurassic of Brunn (Solnhofen Archipelago, southern Germany). PeerJ 9: e11363. DOI: 10.7717/peerj.11363
  65. Voigt O, Fradusco B, Gut C, Kevrekidis C, Vargas S, Wörheide G (2021) Carbonic anhydrases: An ancient tool in calcareous sponge biomineralization. Frontiers in Genetics 12:383. DOI: 10.3389/fgene.2021.624533
  66. Walker C, Harper CJ, Brundrett M & Krings M (2021) The Early Devonian fungus Mycokidstonia sphaerialoides from the Rhynie chert is a member of the Ambisporaceae (Glomeromycota, Archaeosporales), not an ascomycete. Review of Palaeobotany and Palynology 287:104384. DOI: